
Reports & Publications
Peer Reviewed Scientific Publications
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Khan A, Patel K, Shukla H, Viswanathan A, van der Valk T, Borthakur U, Nigam P, Zachariah A, Jhala YV, Kardos M, Ramakrishnan U. Genomic evidence for inbreeding depression and purging of deleterious genetic variation in Indian tigers. Proceedings of the National Academy of Sciences Dec 2021, 118 (49). e2023018118; DOI:10.1073/pnas.2023018118.
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Armstrong E, Khan A, Taylor RW, Gouy A, Greenbaum G, Thiéry A. Kang J, Redondo S, Prost S, Barsh G, Kaelin C, Phalke S, Chugani A, Gilbert M, Miquelle D, Zachariah A, Borthakur U, Reddy A, Louis E, Ramakrishnan U. 2021. Recent evolutionary history of tigers highlights contrasting roles of genetic drift and selection. Molecular biology and evolution. 10.1093/molbev/msab032.
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Ghosh-Harihar M, An R, Athreya R, Borthakur U, Chanchani P, Chetry D, Datta A, Harihar A, Karanth KK, Mariyam D, Mohan D, Onial M, Ramakrishnan U, Robin VV, Saxena A, Shahabuddin, Thatte P, Vijay V, Wacker K, Mathur VB, Pimm SL, Price TD. 2019. Protected area and biodiversity conservation in India. Biological Conservation, 237: 114-124.
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Natesh M, Atla G, Nigam P, Jhala YV, Zachariah A, Borthakur U, Ramakrishnan U. 2017. Conservation priorities for endangered Indian tigers through a genomic lens. Nature Scientific Reports, 7:9614. DOI:10.1038/s41598-017-09748-3.
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Borthakur U, Das PK, Talukdar A, Talukdar BK. 2016. Noninvasive genetic census of greater one-horned rhinos in Gorumara National Park, India – A pilot study for population estimation. Oryx, available on CJO2016. doi:10.1017/S0030605314001161.
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Chetry D, Borthakur U, Das RK. 2015. A short note on a first distribution record of White-cheeked Macaque Macaca leucogenys from India. Asian Primates Journal. 5(1): 45-47.
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Das PK, Borthakur U, Sarma H, Talukdar BK. 2015. Population genetic assessment of extant populations of greater one-horned rhinoceros (Rhinoceros unicornis) in India. European Journal of Wildlife Research. DOI 10.1007/s10344-015-0960-2
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Borthakur U, Saini RP, Sen Gupta S, Jakher R, Das C, Das AK, Sarma PK, Talukdar BK, Bharali R. 2013. Noninvasive genetic assessment of population status of tigers (Panthera tigris tigris) in Buxa Tiger Reserve, West Bengal, India. Int. J. Biodiv. & Conserv. 5(1), 27-32.
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Borthakur U, Barman RD, Lahkar BP, Das C, Das PK, Sarma PK, Bhobora C, Swargowari A and Basumatary GC. 2013. Use of noninvasive genetic tools in monitoring tigers in situ: a preliminary study from Manas National Park, Assam, India. Newsletter and Journal of The Rhino Foundation for nature in NE India 9: 39- 43.
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Tietze DT, Borthakur U. 2012. Historical biogeography of tits (Aves: Paridae, Remizidae). Organisms, Diversity & Evolution, DOI: 10.1007/s13127-012-0101-7.
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Borthakur U, Barman RD, Das C, Basumatary A, Talukdar A, Ahmed MF, Talukdar BK, Bharali R. 2011. Noninvasive genetic monitoring of tiger (Panthera tigris tigris) population of Orang National Park in the Brahmaputra floodplain, Assam, India. European Journal of Wildlife Research. 57 (3), 603-613.
Non-Peer Reviewed Scientific Publications
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Borthakur U, Das C. (2010). "First sighting record of Hooded Pitta, Pitta sordida from Kaziranga National Park, India". Indian Birds, 5 (5): 158.
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Borthakur U. (2009) Noninvasive population genetic monitoring of Indian rhino in Assam: A pilot study to standardize protocols for dung DNA analysis. Rhino Print (Newsletter of the Asian Rhino Project) no. 4, (March), 6-7.
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Borthakur U, Dutta J, Talukdar B. (2008). Noninvasive population genetic monitoring of Indian rhinos in Assam. The Rhino Print (Newsletter of the Asian Rhino Project) no. 1 (August), 4-5.
Technical Reports & Manuals
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Borthakur U (ed.). 2025. Services & Sample Collection Guideline on Wildlife DNA Forensic Investigations. Wildlife Genetics Laboratory, Aaranyak. 1-29 p.
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Saikia A, Sarkar A, Borthakur U, Abedin I, Das AK, Boruah A, Gogoi NK, Das PK, Das BJ, Gayary R, & Lahkar BP. 2025. Understanding the landscape-level corridor connectivity of Asian elephants in Meghalaya (Technical report, 70 pp.). Submitted to Meghalaya Basin Development Authority.
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Borthakur U, Das PK, Deb M, Maheen R, Neog A, Biswas A, Das AK and Barua B (2021). Leopard (Panthera pardus) ecologyin North Bengal including population estimation, distribution, habitat and use pattern and human-leopard conflicts. Aaranyak,Technical Report, Pg. 1-66.
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Borthakur U, Das PK, Deb M, Maheen R, Neog A, Das AK and Barua B (2019). Population status of tiger in North Bengal. Aaranyak, Technical Report, Pg. 1-63.
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Borthakur U, Chetry D, Das RK. 2015. A Rapid faunal survey of Anjaw district, Arunachal Pradesh. Aaranyak Report, Pg. 1-19.
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Borthakur U, Das PK. 2014. Noninvasive genetic monitoring of tiger in Tiger Reserves of Arunachal Pradesh - Survey in Namdapha and Pakke Tiger Reserve. WGP, Aaranyak, Technical Report, Pg. 1 - 25.
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Borthakur U, Das PK. 2014. Genetic analysis based individual identification of snow leopards in Wangchuck Centennial Park, Bhutan. WGP, Aaranyak, Technical Report, Pg. 1 - 16.
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Borthakur U, Das PK. 2014. Assessment of mitochondrial genetic diversity of White-bellied Heron in Bhutan. WGP, Aaranyak, Technical Report, Pg. 1 -11.
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Borthakur U, Das PK. 2014. Ushering Tiger Conservation in Dampa Tiger Reserve, Mizoram, India through Genetic Population Monitoring. WGP, Aaranyak, Technical Report, Pg. 1 - 23.
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Borthakur U, Barman RD, Lahkar BP, Basumatary GC, Swargowari A, Bhobora C, Sarma PK, Das A, Brahma N, Talukdar BK (2013). Noninvasive Genetics Identity Based Monitoring of Tiger in Manas National Park, India. Technical report, Aaranyak WGP, pp. 1 – 20.
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Borthakur U, Das PK, Talukdar BK. 2012. Population genetic monitoring of Greater One-Horned Rhino (Rhinoceros unicornis) in Gorumara National Park, West Bengal, India. WGP, Aaranyak, Technical report no. 9, pp. 1 – 18.
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Borthakur U, Barman RD, Borah J, Puia L, Sarmah A, Vattakavan J, (2012). Monitoring tiger presence in Dampa Tiger Reserve through genetic analysis of scat samples. Technical report, pp. 1 – 10.
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Borthakur U, Barman RD (2012). Genetic identification individual tigers in Namdapha Tiger Reserve, Arunachal Pradesh. Technical report, Aaranyak WGP, No. 10, pp. 1 – 5.
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Borthakur U, Barman RD (2011). Genetic identification of individual tigers in Palamau Tiger Reserve from their scats. Technical report, Aaranyak WGP, pp. 1 – 11.
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Borthakur U, Barman RD (2011). Genetic identification of species identity of carnivore scat samples from Namdapha Tiger Reserve, Arunachal Pradesh. Technical report, Aaranyak WGP, pp. 1 – 6.
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Borthakur U, Basumatary GC, Swargowari A, Bhobora C, Das C, Barman RD, Das PK, Sarma PK, Brahma N, Lahkar BP, Talukdar BK. 2011. Preliminary noninvasive genetic assessment of the status of tigers in Manas Tiger Reserve, Assam. Technical report, Aaranyak WGP, pp. 1 – 17.
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Borthakur U, Das C, Talukdar B K. 2010. Genetic assessment of the status of tigers in Buxa Tiger Reserve, West Bengal. Technical Report, Aaranyak, WGP: 02/2010, pp. 1 – 19.
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Borthakur U, Dutta J, Talukdar BK. 2009. Noninvasive population genetic monitoring of greater one horned rhino (Rhinoceros unicornis) in Assam: a pilot study to standardize protocols for dung DNA analysis. Technical Report_Aaranyak_WGP_01/2009.pp 1- 21.